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Publications
Manuscripts Under Review & In Revision
Berrian, N., Keller, A. S., Chao, A. F., Stier, A. J., Moore, T. M., Barzilay, R., Berman, M. G., Kardan, O. &
Rosenberg, M. D. The exposome and attention-related brain networks jointly predict attention problems in early
adolescence. medRXiv. Under Review.
Lichenstein, S. D., Weng, Y., Robinson, H.+, Rodriguez, L., Babaeianjelodar, M., Maynard, J., Metayer, M., Suneja,
S., Horien, C., Greene, A. S., Constable, R. T., Moore, T. M., Barzilay, R., Yip, S. W., Keller, A. S. Multivariate
environmental exposures are reflected in whole-brain functional connectivity and cognition in youth. bioRXiv. Under
Review.
Nakua, H., Korbmacher, M., Keller, A. S., Keyes, K., Jacobs, G., Wolfers, T., Caye, A., Lee, S. & Marsh, R.
Improving conceptual credibility in developmental neuroimaging research: considerations and strategies to guide
cumulative progress. Under Review.
Juarez, D., Jennings, K., Keller, A. S. & Miller, J. Prediction versus explanation of adolescents'
psychopathology: environmental data predict while brain measures moderate effects. In Revision.
Schneider, D., Bouhali, F., Keller, A. S., Xia, Z., Clement-Lam, S., O’Malley, J., McDonald, C., Luo, J., Gu, C.,
Hoeft, F. Letters from Home: Sex-Discordant Patterns of Parent-Child Correspondence in Reading Achievement.
Under Review.
Xu, X., Fu, Z., Xu, H., Zhang, K., Yang, H., Li, Y., Zhao, Y., Zheng, X., Keller, A. S., Xu, T., Chen, R., Cai, W., Cao,
Q., Yang, L. & Cui, Z. Developmental deviations of structural connectivity in youths with ADHD predict symptom
and treatment outcomes. bioRXiv. Under Review.
Baten, C., Keller, A. S., Miller, C. H. & Sacchet, M. D. The Functional Neuroimaging of Meditation: A Quantitative
Whole-Brain Meta-Analysis and Systematic Review. In Revision.
Ehmann, S., Potash, R., Lautz, N., Baten, C., Keller, A. S.* & Sacchet, M. D.* Cognitive science, attention, and
advanced meditation: Understanding the potential of the human mind. Under Review.
2026
Ojha, A. & Keller, A. S. (2026). From deficits to strengths: how individual differences in neurodevelopment may
reflect adaptations to environmental context and foster resilience. Biological Psychiatry: Cognitive Neuro- science
and Neuroimaging.
Robinson, H., Dave, N., Barzilay, R., Wagner, A., Kells, N. & Keller, A.S. (2026). The effect of the exposome on
developmental brain health and cognition. Neuropsychopharmacology.
Baten, C., Zamora, G., Klassen, A. M., Rackley, J., Nimarko, A. F., Woo, E., Keller, A.S., Hamilton, J. P., Hedges,
D. W., Sacchet, M. D., Gotlib, I. H. & Miller, C. H. (2026). Major depressive disorder in youth and adults: A
quantitative whole- brain meta-analysis of functional magnetic resonance imaging studies. Biological Psychiatry:
Cognitive Neuroscience and Neuroimaging.
Warthen, K. G., Keller, A. S. & Williams, L. M. (2026). Relationship of behavioral drive to time-varying sub-network coherence in attention and reward neural networks. Brain Topography.
Hillman, N., Weinstein, S. M., Bagautdinova, J., Sun, K. Y., Cieslak, M., Salo, T., Fan, Y., Keller, A. S., Alexander- Bloch, A. F., Vandekar, S. N., Raznahan, A., Satterthwaite, T. D., Shou, H. & Shinohara, R. T. (2026). Testing for network specificity in brain-behavior associations using ordinal dominance curves. Human Brain Mapping.
2025
Keller, A. S., Shetty, A., Barzilay, R., Calkins, M. E., Chong, Y.-S., Fair, D. A., Gluckman, P., Gur, R. E., Gur, R. C., Mackey, A. P., Meaney, M. J., Moore, L. A., Moore, T. M., Satterthwaite, T. D., Tan, A. P., Tervo-Clemmens, B. & Larsen, B. (2025). School’s out for the summer: cognition varies across the calendar year in multiple large-scale datasets. Proceedings of the National Academy of Sciences (PNAS).
Ehmann, S., Sezer, I., Keller, A. S., Treves, I. N. & Sacchet, M. D. (2025). Attention and meditative development: a
review and synthesis of long-term meditators and outlook for the study of advanced meditation. NeuroImage
Luo, A. C., Meisler, S. L., Sydnor, V. J., Alexander-Bloch, A., Bagautdinova, J., Barch, D. M., Bassett, D. S.,
Davatzikos, C., Franco, A. R., Goldsmith, J., Gur, R. E., Gur, R. C., Hu, F., Jaskir, M., Kiar, G., Keller, A. S., Larsen, B., Mackey, A. P., Milham, M. P., Roalf, D. R., Shafiei, G., Shinohara, R. T., Somerville, L. H.,
Weinstein, S. M., Yeatman, J. D., Cieslak, M., Rokem, A. & Satterthwaite, T. D. (2025). Two axes of white matter development. Nature Communications.
Damme, K., Seidlitz, J. & Keller, A.S. (2025). Neurodevelopmental timing links stress and psychopathology. JAMA
Pediatrics.
Keller, A. S., Sun, K. Y., Francisco, A., Robinson, H., Beydler, E., Bassett, D. S., Cieslak, M., Cui, Z., Davatzikos, C., Fan, Y., Gardner, M., Kishton, R., Kornfield, S. L., Larsen, B., Li, H., Linder, I., Pines, A., Pritschet, L.,
Raznahan, A., Roalf, D. R., Seidlitz, J., Shafiei, G., Shinohara, R. T., Wolf, D. H., Alexander-Bloch, A.,
Satterthwaite, T. D. & Shanmugan, S. (2025). Reproducible Biological Sex Differences in Personalized
Functional Network Topography in Youth. British Journal of Psychiatry
Sun, K. Y., Schmitt, E., Moore, T. M., Barzilay, R., Almasy, L., Schultz, L. M., Mackey, A. P., Kafadar, E., Sha, Z.,
Seidlitz, J., Mallard, T. T., Cui, Z., Li, H., Fan, Y., Fair, D. A., Satterthwaite, T. D., Keller, A. S.* & Alexander- Bloch, A.* (2025). Polygenic Risk Underlies Overall Psychopathology and Personalized Functional Brain
Network Topography in ABCD. JAMA Psychiatry
Lynn, A. & Keller, A. S. (2025). Attention Emerges from Regional Neural Computations and Selective Information
Flow. In The Oxford Handbook of Infant Attention, Memory, and Learning Development. In Press.
2024
Kang, K., Seidlitz, J., Bethlehem, R. A. I., Mehta, K., Keller, A. S., Schildcrout, J., Tao, R., Xiong, J., Jones, M. T.,
Lifespan Brain Chart Consortium, 3R-BRAIN, AIBL, Alzheimer’s Disease Neuroimaging Initiative, Alzheimer’s
Disease Reposi- tory Without Borders Investigators, CALM Team, CCNP, COBRE, cVEDA, Harvard Brain Aging Study, IMAGEN, POND, The PREVENT-AD Research Group, Fair, D., Satterthwaite, T. D., Alexander-Bloch, A. & Vandekar, S. (2024). Study design features that improve effect sizes in brain-wide association studies. Nature.
Mehta, K.*, Salo, T.*, Madison, T., Adebimpe, A., Bertolero, M., Cieslak, M., Covitz, S., Feczko, E., Houghton,
A., Keller, A. S., Luo, A., Miranda-Dominguez, O., Nelson, S. M., Shafiei, G., Shanmugan, S., Sydnor, V. J.,
Fair, D. A.* & Satterthwaite, T. D.* (2024). XCP-D: Robust Pipeline for the postprocessing of fMRI data. Imaging
Neuroscience.
Zhao, S., Su, H., Cong, J., Wen, X., Yang, H., Chen, P., Wu, G., Fan, Q., Ma, Y., Xu, X., Hu, C., Li, H., Keller,
A. S., Pines, A., Chen, R., Cui, Z. (2024). Hierarchical individual variation and socioeconomic impact on
personalized functional network topography in children. BMC Medicine.
Pines, A., Tozzi, L., Bertrand, C., Keller, A. S., Zhang, X., Whitfield-Gabrieli, S., Hastie, T., Larsen, B., Leikauf, J. &
Williams, L. M. (2024). Intergenerational evidence for concurrent, opposed relationships between cognition and psychopathology in youth: implications for impaired daily functions and clinically unenriched studies. JAMA
Psychiatry.
Keller, A. S., Moore, T. M., Luo, A., Visoki, E., Gataviņš, M. M., Cui, Z., Fan, Y., Feczko, E., Houghton, A., Li, H.,
Mackey, A. P., Miranda-Dominguez, O., Pines, A., Shetty, A., Shinohara, R. T., Sun, K. Y., Fair, D. A.*,
Satterthwaite, T. D.*, & Barzilay, R.* (2024). A general exposome factor explains individual differences in
functional brain network topography and cognition in youth. Developmental Cognitive Neuroscience.
Shafiei, G., Keller, A. S.., Bertolero, M., Shanmugan, S., Bassett, D. S., Chen, A. A., Covitz, S., Houghton, A., Luo,
A., Mehta, K., Salo, T., Shinohara, R. T., Fair, D. A., Hallquist, M. N. & Satterthwaite, T. D. (2024). Linking
functional connectivity to symptoms of borderline personality disorder in youth. Biological Psychiatry.
Luo, A., Sydnor, V. J., Pines, A., Larsen, B., Alexander-Bloch, A. F., Cieslak, M., Covitz, S., Chen, A., Esper, N. B.,
Feczko, E., Franco, A. R., Gur, R. E., Gur, R. C., Houghton, A., Hu, F., Keller, A. S., Kiar, G., Mehta, K., Salum,
G. A., Tapera, T., Xu, T., Zhao, C., Fair, D. A., Salo, T., Shinohara, R. T., Milham, M. P. & Satterthwaite, T. D.
(2024). Functional Connectivity Development along the Sensorimotor-Association Axis Enhances the Cortical
Hierarchy. Nature Communications.
2023
Keller, A. S., Pines, A. R., Shanmugan, S., Sydnor, V. J., Cui, Z., Bertolero, M. A., Barzilay, R., Alexander-Bloch, A. F., Byington, N., Chen, A., Conan, G. M., Davatazikos, C., Feczko, E., Hendrickson, T., Houghton, A., Larsen, B., Li, H., Miranda-Dominguez, O., Roalf, D. R., Perrone, A., Shanmugan, S., Shetty, A., Shinohara, R. T., Fan, Y., Fair, D.A.* & Satterthwaite, T. D.* (2023). Personalized functional brain network topography is associated with individual differences in youth cognition. Nature Communications. https://doi.org/10.1038/s41467-
023-44087-0
Shetty, A., Keller, A. S. (2023). Mapping Individual-Specific Regions of the Multiple Demand Network During
Development Reveals Increased Selectivity Associated with Executive Function. The Journal of Neuroscience.
https://doi.org/10.1523/JNEUROSCI.0849-23.2023
Larsen, B., Sydnor, V. J., Keller, A. S., Yeo, B. T. T., & Satterthwaite, T. D. (2023) A critical period plasticity model for the sensorimotor-association axis of neurodevelopment. Trends in Neurosciences.
https://doi.org/10.1016/j.tins.2023.07.007
Keller, A. S.*, Sydnor, V. J.*, Pines, A., Fair, D. A., Bassett, D. S. & Satterthwaite, T. D. (2023). Hierarchical functional system development supports executive function. Trends in Cognitive Sciences.
https://doi.org/10.1016/j.tics.2022.11.005
Pines, A., Keller, A. S., Larsen, B., Bertolero, M., Ashourvan, A., Bassett, D. S., Cieslak, M., Covitz, S., Fan, Y.,
Feczko, E., Houghton, A., Rueter, A. R., Tapera, T., Vogel, J., Weinstein, S. M., Shinohara, R. T., Fair, D. &
Satterthwaite, T. D. (2023). Development of top-down cortical propagations in youth. Neuron.
https://doi.org/10.1016/j.neuron.2023.01.014
2022​
Keller, A. S., Mackey, A. P., Pines, A., Fair, D., Feczko, E., Hoffman, M. S., Salum, G. A., Barzilay, R. &
Satterthwaite, T. D. (2022). Caregiver monitoring, but not caregiver warmth, is associated with cognition in two large sub-samples of youth from the ABCD study. Developmental Science. https://doi.org/10.1111/desc.13337
Keller, A. S.*, Jagadeesh, A.*, Bugatus, L., Williams, L. M. & Grill-Spector, K. (2022). Attention enhances category
representations across the brain with strengthened residual correlations to ventral temporal cortex. NeuroImage.
https://doi.org/10.1016/j.neuroimage.2022.118900
Richie-Halford, A., Cieslak, M., Ai, L., Caffara, S., Covitz, S., Franco, A. R., Karipidis, I. I., Kruper, J., Milham, M.,
Avelar-Pereira, B., Roy, E., Sydnor, V. J., Yeatman, J. D., The Fibr Community Science Consortium,
Satterthwaite, T. D. & Rokem, A. (2022). An analysis-ready and quality controlled resource for pediatric brain
white-matter research. Scientific Data. https://doi.org/10.1038/s41597-022-01695-7
2021
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Holt-Gosselin, B., Keller, A. S., Chesnut, M., Ling, R., Grisanzio, K. A., Williams, L. M. (2021). Greater baseline
connectivity of the salience and negative affect circuits are associated with natural improvements in anxiety over
time in untreated participants. Journal of Affective Disorders. https://doi.org/10.1016/j.jad.2021.08.039
Keller, A. S., Ling, R., Williams, L. M. (2021) Spatial attention impairments are characterized by specific electro-
encephalographic correlates and partially mediate the association between early life stress and
anxiety. Cognitive, Affective and Behavioral Neuroscience. https://doi.org/10.3758/s13415-021-00963-0
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Goldstein-Piekarski, A. N.*, Ball, T. M.*, Samara, Z., Staveland, B. R., Keller, A. S., Fleming, S. L., Grisanzio, K.
A., Holt-Gosselin, B., Williams, L. M. (2021). Mapping neural circuit biotypes to symptoms and behavioral
dimensions of depression and anxiety. Biological Psychiatry.
2020
Keller, A. S., Davidesco, I., & Tanner, K. D. (2020). Attention Matters: How orchestrating attention relates to
classroom learning. Cell Biology Education - Life Sciences Ed. 19(3):fe5.
https://doi.org/10.1187/cbe.20-05-0106
Rajpurkar, P., Dass D., Yang J., Vale, V., Keller, A. S, Irvin, J., Taylor, Z., Basu, S., Ng, A. & Williams, L. M. (2020).
Machine Learning Prediction of Treatment Response to Antidepressant Medication Using Pre-Treatment EEG: Development and Validation of the ElecTreeScore Algorithm, JAMA Network Open, https://doi.org/10.1001/ jamanetworkopen.2020.6653
Chilver, M. R., Keller, A. S., Park, H., Jamshidi, J., Montalto, A., Schofield, P. R., Clark, C. R., Harmon-
Jones, E., Williams, L. M.*, Gatt, J. M.* (2020). Electroencephalography power as a biomarker of wellbeing: A twin study. Journal of Psychiatric Research. https://doi.org/10.1016/j.jpsychires.2020.04.010
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2019​
Keller, A. S., Ball, T. M. & Williams, L. M. (2019). Deep phenotyping of attention impairments and the
“Inattention Biotype” in Major Depressive Disorder. Psychological Medicine. 1-10. https://doi.org/10.1017/S0033291719002290.
Keller, A. S., Leikauf, J. E., Holt-Gosselin, B., Staveland, B. R. & Williams, L. M. (2019). Paying Attention
To Attention in Depression Translational Psychiatry. https://doi.org/10.1038/s41398-019-0616-1
Keller, A. S., Qiu, H.*, Li, J.* & Williams, L. M. (2019). Modeling attention impairments in Major
Depression. Proceedings of the Computational Cognitive Neuroscience Conference. https://doi.org/10.32470/CCN.2019.1325-0
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2017​
Keller, A. S. & Christopher, L. (2017). Distinct Phases of Tau, Amyloid, and Functional Connectivity in
Healthy Older Adults. The Journal of Neuroscience. https://doi.org/10.1523/JNEUROSCI.1687-17.2017
Keller, A. S., Payne, L. & Sekuler, R. (2017) Characterizing the roles of alpha and theta oscillations in
multisensory attention. Neuropsychologia. https://doi.org/10.1016/j.neuropsychologia.2017.02.021
2015​
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Keller, A. S. & Sekuler, R. (2015). Memory and learning with rapid audiovisual sequences. Journal of Vision.
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